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Comparative anatomy & physiology leave no doubt that human ancestors spent a lot of time diving for shallow-aquatic prey such as shellfish (cf. the so-called "aquatic ape hypothesis" of Alister Hardy & Elaine Morgan). Detailed comparisons suggest:
-Mio-Pliocene hominoids (fossil apes, incl. Plio-Pleistocene australopithecines) evolved: larger size, very broad sternum (Latisternalia), wide thorax & pelvis, centrally-placed spine (more orthogrady), longer arms, more plantigrady, tail loss (coccyx incorporated into pelvic bottom). All this can best be explained by an aquarboreal lifestyle (aqua=water, arbor=tree), wading bipedally & climbing arms overhead in the branches above in flooded forests, e.g. mangroves, forest swamps etc. Australopiths were no closer relatives of Homo than of Pan or Gorilla, google "aquarboreal ancestors". Extant apes apparently survived the Pleistocene coolings by spending less time in the water, but google e.g. "bonobo wading" or "gorilla bai".
-Early-Pleistocene Homo evolved a littoral lifestyle, frequently diving for shallow-aquatic foods such as shellfish: this best explains Homo's brain enlargement, pachy-osteo-sclerosis, platycephaly, external nose (google "Oi big nose"), full plantigrady, platypelloidy, platymeria (dorso-ventrally flattened femora), stone tool use & island colonizations (e.g. Flores), google "coastal dispersal Pleistocene Homo PPT Verhaegen". From the coasts, different "archaic" Homo populations, such as neandertals, entered the land probably initially seasonally along the rivers. Late-Pleistocene H.sapiens reduced diving, and elaborated wading-walking, cf. diet of e.g. rice etc.
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Marc Verhaegen